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the muscles, by being directed to the segment of the cord whence
their nerves proceed. All muscular movements, whether reflex,
emotional, or voluntary, are immediately called into action by
the same class of sensory or efferent nerve fibres ; moreover, the
excitor or afferent fibres are at once iu like manner the chan-
nels of the transmission of stimuli, which gives rise, in one
instance, to reflex actions through the spinal cord, and of im-
pressions, in another instance, that become sensations when
transmitted to the sensorium. The mechanism, then, of a vol-
untary action in these parts that are supplied by spinal nerves
would be the following: —
The impulse of volition, excited primarily in the brain, acts
at the same time upon the gray matter of the cord, (through
the fibres of the white substance of the brain, which radiate
from the thalami optici and the corpora striata, or, according to
Dr. Todd, simply through the strands of the medulla oblongata,
especially the anterior pyramids,) and then from this gray mat-
ter of the cord, simultaneously upon the anterior roots of the
nerves implanted in it. This gray matter, in virtue of its asso-
ciation with the brain by means of the fibres of the white sub-
stance of the brain, or the anterior pyramids, becomes part and
parcel of the organ of the " will" and therefore as distinctly
amenable to acts of the mind, as that portion which is contained
within the cranium. If we destroy the commissural connection
with the brain through the pyramidal or white brain fibres, the
spinal cord ceases to take part in or respond to mental nervous
actions ; or, if that connection be only partially destroyed, then
precisely that portion of the cord, which the now injured nerve
fibres had formerly associated with the brain, is no longer influ-
enced by the mind.
Again, if the seat of volition in the brain be diseased, then
the portion of the cord that corresponds to the seat of that
lesion participates in the effects of the disease as far, at
least, as regards voluntary actions. And again, we must bear
in mind that a continuity of fibrous structure here is not
absolutely necessary for the transmission of nervous energy,
or, as Dr. Todd would say, ufor the excitement of nervous