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Galvani, Volta, and Yalli, contended that muscular contrac-
tions were rendered most strong from the immediate contact of
the metallic stimulus to
the cut and raw surface
of the muscle, without
touching the nerve. But
others, as Baron Humboldt
and Dr. Fowler, thought
that such test could never
si give a fair solution of the
sf question since it is almost
impossible to eradicate all
nerve fibrils from muscle
fibres. They, therefore,
were led to conclude that
irritability is a property of
nerve and muscle as a compound structure.
Dr. Marshall Hall announced the same opinion some fifty
years afterwards.f Whether the property of irritability belongs
to the pure and isolated muscular fibre, or to the muscle fibre
combined with the nerves, he thought, could not be positively de-
termined by distinct experiment; and that irritability belonged,
in all probability, to the compound structure.
The law of Yalli and Ritter, which is a confirmed fact in
pathology, is in truth the greatest hindcrance to determining
this yet unsettled question. That law is, " The nerves may die in
Fig. 49. A View of Elementary Fibres of Voluntary
MuhcIcs; i. e., of animal life, magnified three hundred
diameters, showing a cleavage under slrain, both
lengthwise nnd transverse. This also shows that lame-
ness from strained muscles may result from ruptured
fibres.*
* 1. The longitudinal cleavage, also showing the molecular structure.
2. The transverse cleavage.
3. Incomplete fracture of a muscle fibre, following the opposite surfaces of a
disk, which stretches across the interval, and retains the two fragments in connection.
4. Shows another disk, nearly detached, as often seen.
5. A detached disk of molecules, still more highly magnified, showing the sarcous
element of muscle fibres.
6. Fibrillee separated by the effects of violence from each other at the torn end of
the muscle fibre.
7. 8. The two appearances, commonly observed under the microscope, by the single
Fibrillte. At 7 they are more highly magnified and appear rectangular ; but at 8 the
borders are rounded, and the spaces are bead-like.
t Cyclopaedia of Anatomy and Physiology, vol. iii. p. 29, 1847.